Pokémon HeartGold and SoulSilver are role-playing video games with adventure elements. The basic mechanics of the games are largely the same as their predecessors'. As with all Pokémon games for handheld consoles, gameplay is viewed from a third-person overhead perspective, and consists of three basic screens: a field map, in which the player navigates the main character; a battle screen; and the menu, in which the player configures their party, items, or gameplay settings. The player begins the game with one Pokémon and can capture more using Poké Balls.

When the player encounters a wild Pokémon or is challenged by a trainer to a battle, the screen switches to a turn-based battle screen where the Pokémon fight. During battle, the player may use a move, use an item, switch the active Pokémon, or flee. Fleeing is not an option during battles against trainers. Pokémon have hit points (HP), which are displayed during battles. When a Pokémon's HP is reduced to zero, it faints and cannot battle unless taken to a Pokémon Center or healed or revived with a Pokémon skill or an item, typically a 'revive' medicine. If the player's Pokémon defeats the opposing Pokémon (causes it to faint), it receives experience points. After accumulating enough experience points, it will level up; most Pokémon evolve into a new species of Pokémon when they reach a certain level, or when certain conditions are met, such as how much a Pokémon statistically 'likes' its trainer.

New features of Pokemon SoulSilver:

In HeartGold and SoulSilver, the first Pokémon in the player's party can follow them in the over world, echoing a mechanic in Pokémon Yellow where Pikachu follows the player. This mechanic was also used in a limited fashion in Pokémon Diamond, Pearl, and Platinum when the player is in Amity Park with a "cute" Pokémon. The player may talk to the Pokémon to see or check on how that Pokémon is feeling, and occasionally it may pick up items. Additionally, a pedometer was bundled with each copy of the games called a "Pokéwalker", which allows players to transfer Pokémon from the game into the device and walk around with them, earning "watts" in the process, which can be exchanged for in-game rewards.

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